An executive at Google said it was not aware of the alleged
activity, adding there an "urgent need for reform".
The NSA's director said it had not had access to the
companies' computers.
Gen Keith Alexander told Bloomberg TV: "We are not
authorised to go into a US company's servers and take data."
But correspondents say this is not a direct denial of the
latest claims.
'Extending encryption'
The revelations stem from documents leaked by fugitive ex-US
intelligence contractor Edward Snowden, who now lives in Russia and is wanted
in the US in connection with the unauthorised disclosures.
The documents say millions of records were gleaned daily
from the internet giants' internal networks.
They suggest that the NSA intercepted the data at some point
as it flowed through fibre-optic cables and other network equipment connecting
the companies' data centres, rather than targeting the servers themselves.
The data was intercepted outside the US, the documents
imply.
The data the agency obtained, which ranged from
"metadata' to text, audio and video, were then sifted by an NSA programme
called Muscular, operated with the NSA's British counterpart, GCHQ, the
documents say.
The NSA already has "front-door" access to Google
and Yahoo user accounts through a court-approved programme known as Prism.
Google's chief legal officer, David Drummond, said Google
did not provide any government with access to its systems.
"We have long been concerned about the possibility of
this kind of snooping, which is why we have continued to extend encryption
across more and more Google services and links, especially the links in the
slide," Drummond said in a statement.
"We are outraged at the lengths to which the government
seems to have gone to intercept data from our private fibre networks, and it
underscores the need for urgent reform."
A spokesperson for Yahoo said the company had "strict
controls in place to protect the security of our data centres, and we have not
given access to our data centres to the NSA or to any other government agency".
An NSA spokesperson denied a suggestion in the Washington
Post article that the agency gathered "vast quantities of US persons' data
from this type of collection".
The latest revelations came hours after a German delegation
of intelligence officials arrived in Washington for talks at the White House
following claims that the US monitored Chancellor Angela Merkel's mobile phone.
Two of Mrs Merkel's most important advisers, foreign policy
adviser Christoph Heusgen, and intelligence coordinator Guenter Heiss were sent
to take part in the talks - seen as a measure of how seriously Mrs Merkel takes
the matter.
Next week, the heads of Germany's spying agencies will meet
their opposite numbers in Washington.
'Inappropriate and unacceptable'
The head of US intelligence has defended the monitoring of
foreign leaders as a key goal of operations but the US is facing growing anger
over reports it spied on its allies abroad.
It has also been reported that the NSA monitored French
diplomats in Washington and at the UN, and that it conducted surveillance on
millions of French and Spanish telephone calls, among other operations against
US allies.
Spanish Prime Minister Mariano Rajoy said that if Spain had
been a target of the NSA, this would be "inappropriate and unacceptable
between partners".
However, Gen Alexander has said "the assertions... that
NSA collected tens of millions of phone calls [in Europe] are completely
false".
On Wednesday, the agency denied Italian media reports that
it had targeted communications at the Vatican.
The UN said it had received assurances that its
communications "are not and will not be monitored" by American
intelligence agencies, but refused to clarify whether they had been in the
past.
On Tuesday, Director of National Intelligence James Clapper
testified before the intelligence panel of the House of Representatives that
much of the data cited by non-US news outlets was actually collected by
European intelligence services and later shared with the NSA.
He said foreign allies spied on US officials and
intelligence agencies as a matter of routine.
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